Conceptualization, B.X. and B.Z.; Writing—original draft preparation, S.X., B.X. and B.Z.; Writing—review and editing, X.W. and S.X.; supervision, X.W. and S.X.; funding acquisition, X.W. and S.X. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (22261160370 and 21905006), the Natural Science Foundation of Hunan Province, China (2023JJ50132), the Guangdong Provincial Science and Technology Plan (2021A0505110003), the Guangdong Basic and Applied Basic Research Foundation (2020B1515120039), the Shenzhen Science and Technology Program (JCYJ20220818100211025, ZDSYS20200811143600001 and 2022378670), and the Natural Science Foundation of Top Talent of SZTU (GDRC202343).