Conceptualization, T.M., J.Z., X.L., Q.Z., Z.L. and F.Y.; methodology, T.M., J.Z., X.R., L.Y., S.Z. and F.Y.; validation, T.M. and X.R.; curation analysis, T.M. and J.Z.; writing—original draft, T.M., X.R. and J.Z.; writing—review and editing, F.Y.; funding acquisition, F.Y. All authors have read and agreed to the published version of the manuscript. This work was financially supported by the Key Research and Development Projects in Hunan Province (2022SK2089), the Hunan Provincial Natural Science Foundation (2023JJ40539), and the Huxiang Youth Talent Program (2021RC3107).