* Corresponding authors. E-mail: weiminhe@usc.edu.cn; jiangjun9005@163.com Received March 28, 2022; revised April 26, 2022; published online May 6, 2022. Project supported by the Scientific Research Fund of Hunan Provincial Education Department (No. 19C0783) and the Science and Technology Innovation Program of Hunan Province (No. 2021RC5028). 湖南省教育厅科学研究项目(No. 19C0783)、湖南省科技创新计划项目(No. 2021RC5028)资助项目.