CRediT authorship contribution statement Ximei Sun: Software, Methodology, Data curation, Conceptualization. Feirong Wang: Software, Methodology, Data curation. Jieqiong Wang: Methodology, Formal analysis. Hao Xiao: Methodology, Formal analysis. Qiang Kang: Methodology, Investigation. Yiqian Zeng: Investigation, Data curation. Qinglai Yang: Writing – review & editing, acquisition, Conceptualization. Xiaofeng Tan: Writing – original draft, Supervision, Conceptualization. Acknowledgments Q.Y. acknowledges financial support from the National Natural Science Foundation of China (22374065), the Science and Technology Innovation Program of Hunan Province “Huxiang Young Talents Plan” (2021RC3106), and the Key Research and Development Program of Hunan Province, China (2022SK2053). X.T. thanks the Natural Science Foundation of Hunan Province of China (2022JJ40360) and the Scientific Research Fund of Hunan Province Education Department (22B0409) for funding support. Y.Z. thanks the