J.Z. conceived and wrote the manuscript; J.W., I.Y.M. and T.P. edited the paper. F.Y. conceived and edited the paper and supported funding. All the authors participated in the preparation of the manuscript. All authors have read and agreed to the published version of the manuscript. This work was supported by Huxiang Youth Talent Program (2021RC3107); the Key Research and Development Projects in Hunan Province (2022SK2089); Hunan Province Excellent Youth Fund (2020JJ3053); National Natural Science Foundation of China (81773393, 31900108); the Scientific Research Start-up Fund of University of South China (201RGC008), Natural Science Foundation of Hunan Province of China (2022JJ40372).